Is an ortholog of human CARMIL1 (capping protein regulator and myosin 1 linker 1) and CARMIL3 (capping protein regulator and myosin 1 linker 3). Exhibits GTPase activating protein binding activity. Is involved in generation of neurons\; negative regulation of Rac protein signal transduction\; and regulation of cell migration. Localizes to the plasma membrane. Is expressed in the alimentary system\; the epithelial system\; the excretory canal\; the nervous system\; and the reproductive system.